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Fixed the overlapping UVs but mesh has still parts which are 100% dark

I had a warning with overlapping UVs on a foliage mesh and so I fixed the overlapping UVs so the warning doesn’t come after building but the mesh has still parts which are 100% dark.
What can I do? Did I something wrong?

https://picload.org/image/rardrplo/zwischenablage00.jpg

https://picload.org/image/rardrplc/zwischenablage01.jpg

https://picload.org/image/rardrplp/zwischenablage02.jpg

https://picload.org/image/rardrpcg/zwischenablage03.jpg

https://picload.org/image/rarddllw/zwischenablage04.jpg