I’ve confirmed that for a new 5.3.2 project, changing the mesh renderer to particle was the only change necessary. The project already had heterogeneous volumes turned on, so I also confirmed that turning it off breaks the Niagara system again. Turning this on for the emitter itself was not necessary.
For anyone wondering where this setting is, click the master emitter in the blueprint, search for “mesh renderer” and look for “Source Mode”
