I’ve gotten the lighting setup to work both ways, as long as everything is in the same level.
The black shadows could be caused by distance field shadows.
Select your directional light, then under the Distance Field Shadows section, uncheck “Use RayTraced DistanceField Shadows.”
BTW the default time of day effects when you load a map are not correct, so don’t judge your lighting based off of it. Open your matinee_timeofDay matinee sequence and scrub to a different time.