Howto setup Non-conform Modular Skeletal Meshes at Runtime

Every skeleton has re-target settings on each bone.
they are normally hidden, you have to rigjt click and choose the appropriate option for them to display.
by default the settings are a mixed mess / if you are doing animations and different skeletons they may need to be set to “animation”. If you do use retargeting, then they likely need to be set to skelton retargeting. Really depends on what you need, but it may be a cause for the bone being out of place.