The low-res video is a little hard to see, but here’s what it looks like is going on with the Start transition. In the Start to Locomotion transition, you’re ANDing TimeRemaining with the control logic. Which means that if both MoveRight and MoveForward are 0 while you’re playing the Start animation, Can Enter Transition will always be False, and so the transition will never leave. What’s going to happen then is that your Start animation will complete and freeze at the end (or worst, will restart if you’ve accidentally set your Start animation to loop) until you’ve pressed WASD.
Try doing this:
Also, make sure your Start BlendSpace is set to not loop. This way, the transition will leave when the Start animation has (nearly) completed. If you tap WASD, the Start animation will play all the way through, and then you’ll transition to your Locomotion state.