I had bad luck of reproducing this error in another project. UE4 crashed (together with Oculus and GPU driver).
When I turned project on, my EnemyMelee which is child of Enemy has started to loosing mesh, animation and location every time i turn the project on again.
I’m not sure if the crashed was the case or just a coincident.
I have actor that spawns enemies. If I remove EnemyMelee from spawning, then it’s not loosing values on restart.