After some time, I have finally come back to this to do some more debugging. It turns out, some of my animations are causing this issue on specific levels. Why is it level specific, I have no idea. I did find, if I change my Idle animation, which I got from Mixamo and converted, to an Idle animation from the started pack this starts working again. I am guessing this has to do with some odd root motion issue? Again I am unsure. I may keep banging on this, but ultimately I think I am going to just rebuild my characters from scratch, including the Animation Blueprint, with better converted animations. I am going to use the plugin from the Unreal store and see if I get a different outcome.
Please, if anyone has any suggestions upon this new lieu of evidence, please let me know as this will be a process to re-do all that existing work. Nonetheless, game development is never easy!