Why is my animation playing in Matinee but not PIE?

The Director has no incidence on animations or movements: it only directs which camera feed is the main one at the time. It contains a reference to Camera40 because that’s a Camera Actor. It doesn’t contain a reference to Camera4 because that’s a Skeletal Mesh to which Camera40 is attached. This is the setup I received from animation.

Camera4 and Camera40’s Movement tracks are mostly empty because all I want is to make sure they are at 0;0;0 so that all other animations blend smoothly and it goes correctly with the environment.

The Movement track is NOT what animates Camera4! It is its Animation track, which moves Camera4 correctly in Matinee but not in PIE.