Character off by 90 degrees

Recently the player character started being off by 90 degrees for movement. Simply swapping out the character and the controller didn’t fix it.

Opening the character, then closing it then running the game in Editor fixes the problem.

The packaged game, the character is off by 90 degrees.

I have tried setting the character’s yaw to -90 which will work as long as the Editor is opened. Saving, closing the editor then reopening it, the character’s movement is off again by 90 degrees.

This is version 4.10.2.